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Best linear unbiased estimators (BLUEs) across experiments were used as phenotypes for genome-wide association studies (GWAS). GWAS was conducted for 454 whole-genome sequenced (WGS) wheat genotypes considering a kinship matrix computed from genotyping-by-sequencing (GBS) markers. SNP-based GWAS was conducted using WGS variants mapped to Chinese Spring (RefSeq v1.0) while k-mer-based GWAS was performed using the presence/absence of 31-bp-long sequence motifs. 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